Department of Agriculture, Ehime University, Tarumi, Matsuyama, Ehime 790-8566, Japan. Division of Molecular and Cellular Biology, Graduate School of Bioagricultural Sciences, Nagoya University, Furo-cho, Chikusa, Nagoya, Aichi 464-8601, Japan. Institution for Gene Research, Advanced Science Research Centre, Kanazawa University, 13-1 Takara-machi, Kanazawa, Ishikawa 920-0934, Japan.